Sean Craig

Professor - Zoology and Marine Biology

(707) 826-3656
SciA 457

Research: Population dynamics, ecology and life-history strategies of colonial marine organisms.


BA (1983) University of New Hampshire
MS (1987) University of Houston
PhD (1995) SUNY Stony Brook

Courses Taught

General Zoology
Invertebrate Zoology
Intertidal Ecology
Graduate seminar on competition & cooperation


Research focuses on 2 areas of marine ecology: 1) self-nonself recognition and the evolution of cooperation in colonial marine animals, and (2) rocky intertidal and subtidal community ecology, with a particular focus on interactions between native and introduced species.


  • Pemberton, A. J.,L.J. Hansson, S.F. Craig, R.N. Hughes and J.D.D. Bishop.  2007.  Microscale genetic differentiation in a sessile invertebrate with cloned larvae: investigating the role of polyembryony.  Marine Biology 153 (1):  71-82.
  • Boyle, M., D. Janiak, and S.F. Craig.  2007.  Succession in a Humboldt bay marine fouling community: the role of exotic species, larval settlement and winter storms.  Proceedings of the 2004 Humboldt Bay Symposium published by California SeaGrant.  pgs 215-234. PDF
  • Hughes, R.N., P.H. Manriquez, S. Morley, S.F.Craig, and J.D.D. Bishop.  2005.  Kin or self recognition?  Colonial fusibility of the bryozoan Celleporella hyalina.  Evolution & Development 6 (6):  431-437. PDF
  • Hughes, R.N., M.E. D'Amato, J.D.D. Bishop, G.R. Carvalho, S.F. Craig, L.J. Hansson, M.A.Harley, and A.J. Pemberton.  2005.  Paradoxical polyembryony?  Embryonic cloning in an ancient order of marine bryozoans.  Biology Letters 1: 178-180. PDF
  • (2002) Craig, S.F. Introduced species and their role in marine "fouling" communities in Humboldt Bay. U.S. Fish & Wildlife Service Report. 10pp.
  • (2001) Craig, S. F., D'Amato, M. E., Harley, M., Bishop, J., Hughes, R. & Carvalho, G.R. Isolation and characterization of microsatellites in the bryozoan Crisia denticulata. Molecular Ecology Notes, 100(1): 1-2
  • 2000 Craig, S.F. and Wasson, K. Self/non-self recognition and fusion in the bryozoan Hippodiplosia inscultpa. In P.J Hayward & J. Taylor (eds.). Biology and Paleobiology of Bryozoans: Proceedings of the 11th International Bryozoology Association Conference, Panama City, Panama. pgs 189-196
  • 1997 Craig, S.F., L.B. Slobodkin, and G. Wray. The 'Paradox' of Polyembryony: A review of the cases and a hypothesis for its evolution. Evolutionary Ecology, 11 (2): 127-143.
  • 1997 Judge, M.L. and S.F. Craig. Water flow effects on recruitment and growth of barnacles and hydroids: results from in situ manipulation of water currents. Journal of Experimental Marine Biology & Ecology 210(2): 209-222

Graduate Students

Ismael Chowdhury, Hannah Lee, Jason Lopiccolo, Franklin Moitoza, Dustin Price, Sheena Stephens, Alexander Strawhand, Claire Windecker

Former Students: Johnathan Centoni, Katie Houle, Brendan Kelly, Kellan Korcheck, Michelle Succow

"A long term comparison of rocky intertidal communities in Redwood National and state parks"McGary, Cara L. (Humboldt State UniversityHumboldt State University, 2005-12)

"The effects of temperature on the composition of communities associated with Mytilus californianus". Young, Alison (Humboldt State University, 2009-05)

"Growth inhibition of red abalone (Haliotis rufescens) infested with an endolithic sponge (Cliona sp.)". Morejohn, Kirby (Humboldt State University, 2012-05)

"Growth morphology and succession in a temperate marine fouling community"Nelson, Matthew L. (Humboldt State University, 2009-05)

"High predation may hinder native oyster (Ostrea lurida Carpenter, 1864) restoration in North Humboldt Bay, California"Koeppel, Julie A. (Humboldt State University, 2011-05)

"The facilitative role of an introduced bryozoan (Watersipora spp.): structuring fouling community assemblages within Humboldt Bay"Wilson, Emily Erin (Humboldt State University, 2011-11)

"Distribution, genetic differentiation, and assortative mating of distinct morphotypes of Diaulula sandiegensis, a nudibranch with high dispersal potential"Kelly, Julie A. (Humboldt State University, 2013-05)

"Three factors affecting the invasion of Watersipora subtorquata in the marine fouling community of Humboldt Bay, California"Roscoe, Catherine M. (Humboldt State University, 2005-05)

"Variation in photosynthate release from symbiotic dinoflagellates and its potential role in cnidarian health".(Co-advised by Dr. Ruth Gates, Hawaii Institute of Marine Biology) Morris, Emily (Humboldt State University, 2006-12)

"Boulders over easy: Natural and anthropogenic disturbance in Patrick's Point State Park". Absher, Matt (Humboldt State University, 2004-05)